After the TARDIS makes a crash landing on the Earth of another universe, Rose discovers her father is alive and rich, Mickey encounters his alternative self, and the Tenth Doctor learns one of his oldest and deadliest foes is about to be reborn.

The plans of John Lumic to convert the world into Cybermen are now in full force. The Tenth Doctor, Rose, Mickey and the Preachers must find a way to stop him from enforcing his "ultimate upgrade."

Humanity is caught in the crossfire as the Cybermen and the Daleks wage war against each other. When the mysterious Genesis Ark opens, the Tenth Doctor is faced with an impossible decision: saving the world could mean losing Rose Tyler forever…