Jacks-of-all-trades Yun and Haberu investigate sounds emanating from an empty mansion. They locate the source nearby, while Mei hears the same tune.

A legion of Rodan swarm towards Nigashio. The Otaki Factory crew scramble to lure them away from the city, but soon find themselves surrounded.

Mei discovers a mysterious code in Professor Ashihara's research. The Otaki crew confront Anguirus as part of the city's extermination efforts.

Following a group of Manda, a giant kaiju makes landfall in Tokyo. Red dust forces Mei and Professor Li's flight to London to divert to Milan.

Ground forces under the command of Matsubara respond to Godzilla's attack on Tokyo. Now in London, Mei attempts to decipher Ashihara's research.

Back at Ashihara's mansion, Yun and Haberu attempt to decipher the professor's research using Jet Jaguar. In India, Lina takes Mei to meet with BB.

Mei learns of a large explosion triggered by Ashihara's calculator long ago. Kanoko gives Matsubara the Orthogonal Diagonalizer to use against Godzilla.

Mei, BB and Pelops II attempt to prevent the Catastrophe, but Salunga ambushes them. Meanwhile, the Otaki crew make their final push toward Godzilla.