Will was born in the city of the dead, where he was raised by three undead. Although he was nurtured with love, Will questions his existence in this life-forbidding place where he is the only human alive. He discovers his three parents are secretly hiding something that may explain this mystery, and as if this weren't puzzling enough, he has yet to discover that still more secrets lie within him.

The "God of Undeath" Stagnate's sudden appearance catches the three undead off-guard. Although his attempt to abduct Blood and Mary is interrupted by Gus, the black mist bursting from his chest leaves them no time to feel relieved. Will is unable to do anything as Stagnate says to him with a fearless smile, "It's all thanks to you."

Will and his companions set out for the Rust Mountains to fight the dragon Valacirca. On the way, they pass by the City of the Dead, so Will decides to stop and visit his old home.

Will's crew saves Dine, an elf, from a hydra's attack. Dine tells them that the dragon's miasma has tainted her homeland of Lothdor and sickened its people, so Will decides to go there and heal them.