Shindou Yuu is an amateur model at the top of her class, and Hakozaki Kusue is a physically weak girl who misses school a lot. One moment, Yotsuya Yuusuke is thinking that the sight of these two together seems unusual… and the next, he finds himself in another world. Suddenly Hakozaki, a warrior who can’t even swing a sword, and Yotsuya, whose only weapons are a hoe and a sickle, are a party with zero combat experience stuck with the task of defeating a powerful troll.

The party heads toward Radodorbo, a village far to the west, to complete the latest stage of their quest. However, Shindou is captured by bandits on the way. Tokitate and Hakozaki want to save her, but Yotsuya just wants to hurry on, so the three of them end up parting ways. Yotsuya then enters a martial arts tournament in Cortonel in hopes of winning some horses they can use, and there, he meets a woman knight named Kahvel.

Yotsuya’s party is trapped by the Deokk soldier Kamilto in a cave filled with ancient traps. Yotsuya becomes separated from the group, leaving the other three party members and Kahvel to fight a formidable gargoyle. Things look desperate when the gargoyle takes down Kahvel, but then Shindou reaches rank 10 and job-changes to a Warrior (Sword), allowing her to put up a fight against the gargoyle.

As her reward for completing the quest, Tokitate gets to ask the Game Master whether this alternate world is actually virtual or not... and the answer is that it's an alternate Earth that branched off of the original Earth's timeline. Upon hearing that it's an existing world, Yotsuya trembles with the realization that he killed real people. Once he returns to the real world and resumes his normal life, the Game Master appears before him in the form of a young girl and leads him to a meeting with the next new player.

The boy Yotsuya saved in the Chinese restaurant, Torii Keita, turns out to be the fifth player. They're sent to the alternate world again, and after receiving the fifth quest, they head for Cortonel to track down some leads. There they find Kahvel, who is now a mother. The party is shocked to learn that 15 years have passed since their fourth quest, but that isn't the only thing that's changed in this world.