A padre fresh from Guadalcanal arrives to visit the Black Sheep squadron. Rather than man of courage and faith, they see a man in the middle of a crisis of faith. Meanwhile local Japanese bombers seem to be taking a lot interest in activity around the islands.

Sgt Micklin receives a promotion but he seems very resistant to the idea. Pappy needs him to accept the new position so the chronic issues with ammunition for the Black Sheep squadron can be resolved.

The Black Sheep squadron are in direct competition with an army squadron. The winner of the contest will get the opportunity to fly an extremely dangerous and important mission.

A group of Army pilots arrive at the base. Naturally there is friction between the groups of fliers. Ultimately the friction leads to an opportunity no one can pass up to complete one of the most important missions of the wa

Squadron 214 has to re-qualify for combat duty. Conflict erupts when they discover they will be tested on what the book in Washington says and not direct combat experience against the Japanese.

The island is visited by a number of politicians on a fact finding mission. The pilot flying them is a serious black market operator. And soon horse trading among the men gets out of hand.

A new recruit joins the squadron and the commander has extreme doubts about his age. After a series of encounters it is obvious the young man is an extremely talented flyer and is welcomed into the squadron with open arms.

The base for the squadron comes under attack from Japanese land forces. The situation becomes more desperate with need to evacuate the nursing staff. With all retreat cut of the men and women decide to make their stand.

A friend of Boyington's, who's the number 1 ace, is shot down by a Japanese pilot, known as the Iceman, who only goes after the top pilots. Boyington is now the number one ace pilot and he is told that he has to go back to the States to go on a bond drive. And the one who's going to replace him is not exactly what the squad is use to.

The Japanese are bombing LaCava and they damage the runway. Boyington then asks Lard if he can get some SesBees to repair the runway. Lard tells him if he can find one, he'll authorize it. He does. When he goes to see the man in charge, he says he has too much to do. But when Boyington tells him that a USO show will be visiting the island, he agrees. It seems that while he was at Lard's office, he heard of a USO show coming so he has Casey find out if they can come to the island. But when he calls he is told they are not coming.

The squadron save a transport plane under attack from the Japanese only to find later the wife of the president of the United States was on board. The subsequent publicity becomes overwhelming.

One of the Black sheep has his plane damaged and tries to make for an island known as England. He comes down between Japanese forces and an English coast watcher and the race is on to save him.